Structure and function. Attempting to understand wood in ... bark, vascular cambium, sapwood, heartwood, and the pith (Fig. Developing cambium. vascular cambium. Perspective. The cambium has been variously defined as follows: "The actively dividing layer of cells that lies between, and gives rise to, secondary xylem and phloem (vascular cambium)" (IAWA 1964); "A meristem with products of periclinal divisions commonly contributed in two directions and arranged in radial files. In the following two chapters we shall discuss in detail the structure, functions, and the importance to the plant of these … i.e., secondary xylem and secondary phloem. under Vascular System) are produced every year from a specialized meristem called the vascular cambium (or simply cambium). Structure and Function of Wood Alex Wiedenhoeft, Botanist 3–1 Wood is a complex biological structure, a composite of many chemistries and cell types acting together to serve the needs of a living plant. 5. Interfasicular cambium (from ray parenchyma) fasicular cambium (from procambium) xylem phloem tylose Spring wood Summer wood bark cork. Vascular Cambium The vascular cambium is the lateral meristem that produces the secondary vascular tissues. The location and arrangement of cambium, xylem, and phloem vary between plant parts (e.g., shoots and roots) and with the develop-mental stage of the part. The cambium present between primary xylem and primary phloem is called the intrafascicular cambium (within vascular bundles). LOCATION OF VASCULAR CAMBIUM In Dicots: The vascular cambium is in dicot stems and roots, located between the xylem and the phloem in the stem and root of a vascular plant, and is the source of both the secondary xylem growth (inwards, towards the pith) and the secondary phloem growth (outwards). It is to note that vascular cambium is absent from most monocotyledons, pteridophytes and some herbaceous dicotyledons. Following a general outline on the functioning of the cambium, the authors review the data acquired over the last 20 years. Origin and Formation of Vascular Cambium A strip of vascular cambium that is believed to originate from the procambium is present between xylem and phloem of the vascular bundle. VASCULAR CAMBIUM 4. cortex cortex --> phelloderm cork cambium phellem As the stem enlarges, the epidermis is broken, and must be renewed (as During secondary growth, cells of medullary rays, in a line (as seen in section; in three dimensions, it is a sheet) between neighbouring vascular bundles, become meristematic and form new interfascicular cambium (between vascular bundles). Aristolochia, Clematis etc. 3–1). Wood structure and function change with maturity: Age of the vascular cambium is associated with xylem changes in current‐year growth F. Daniela Rodriguez‐Zaccaro Department of Biology, California State University, Bakersfield, Bakersfield, California It is difficult to overemphasize the importance of the vascular cambium which produces secondary xylem and secondary phloem. tylose Pith rays. Paper IV Structure, Development and Reproduction in Flowering Plants A new pair of papers investigates how the Arabidopsis root vascular cambium forms and how it functions. In woody lianas, e.g. The vascular cambium functions throughout the life of woody perennials. (Fig. The vascular cambium is a meristematic tissue that is responsible for lateral growth and the continued production of new xylem and phloem; in woody plants, the shoot vascular cambium makes wood.